Despite a lack of publications, Leonardo’s fame grew as word of his notebooks spread. The first work by Leonardo to be printed was his Treatise on Painting, published a century after his death. That 1651 Italian edition is on display at the Fred Jones Jr. Museum of Art in Spring 2016. The Treatise on Painting conveys Leonardo’s views on mathematical proportion and the human body, the physiology of vision, and plant physiology. Given Leonardo’s understanding of art as an intellectual endeavor carried out in imitation of nature, it is impossible to separate his art theory from his understanding of science.